Appendix 3.  Photosynthesis Equations Used in CIRAS-2 
 
CIRAS-2 Operator's Manual Version 2.04 
- 159 - 
 
(6.2) Where: 
 
 
 
 
Please note: These calculations are based on the following assumptions: 
 
i)  the leaf is exposed on both upper and lower leaf surfaces 
 
ii)  the upper and lower boundary layer resistances are similar 
 
iii)  stomata are evenly distributed on both upper and lower leaf surfaces. 
 
 
Symbol Definitions 
Measured Parameters 
 
 
Mass flow of dry air into cuvette at STP 
 
Boundary layer resistance to water vapor 
 
Photon flux density incident on cuvette 
 
Calculated Parameters 
 
 
Mass flow of dry air per unit leaf area 
 
Partial pressure of water vapor of air entering cuvette 
 
Partial pressure of water vapor of stirred cuvette air 
 
Saturated vapor pressure at cuvette air temperature 
 
Saturated vapor pressure at leaf temperature, inside the leaf